Karen Hoffman received her Ph.D. in Pharmacology from the Department of Pharmacology and Experimental Neurosciences at the University of Nebraska Medical Center in Omaha, NE, where she was the recipient of an American Heart Association fellowship and several regional and national awards for her research on G protein-coupled receptor signaling in airways. She then pursued post-doctoral research projects at the University of North Carolina-Chapel Hill and the University of Kansas Medical Center, again receiving fellowships from the PhRMA Foundation and the American Heart Association, respectively. She has published research in the American Journal of Pathology, Journal of Biological Chemistry, and Journal of Pharmacology and Experimental Therapeutics. In 2012, Karen joined the editorial staff at WilliamsTown Communications, an editing firm that specializes in educational products for undergraduate- and graduate-level students. At WTC, Karen specializes in producing educational products related to the sciences and healthcare. In addition, Karen is board-certified for editing life sciences (BELS-certified).